bacterial generations - how are they tracked ?

Hi all,

I have been reading a paper by Richared Lenski on a long running
experiment using e coli bacteria.
He says that they have taken a sample and frozen it every 500
generations.
Is that meant literally or an approximation and how do you tell ?

lots of Detail not really required , I dont have the background :( ,
just curious

I found the paper referred to at Wikipedia and then Conservipedia (?)
which seems to be a bit of a whackjob site

regards

.

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